在高级阶元的分子系统发育分析中，充分的取样是一个关键因素，既有助于改善系统发育推论的准确性，也有助于提升所得树图的解释能力。在分子系统发育分析中有两个突出的问题：基部长支的位置不稳定，和高支持度分支的系统发育关系不确定。对于这两个问题，理论和实际研究结果都表明有望通过增加取样来解决。然而，有时分子系统发育与分类系统之间的严重冲突，会导致无论现有的分类系统还是已建立的系统发育，都无法为增加取样提供有效的选择标准。在本研究中我们以蜘蛛外雌器类型为取样依据重建褶蛛-微珠分支高级阶元的系统发育；通过追溯外雌器特征的系统演化，对树图所表现的系统发育关系进行解读；并在系统发育的基础上，对褶蛛的分类系统进行重新修订。我们的研究结果将为在不完全取样的高级阶元系统发育分析中，如何能够获得充分的取样代表性，提供一个可操作的标准；追溯连续变化的外雌器特征的系统演化，为解读有争议的系统发育关系提供一个可行的办法。

For high-level molecular phylogenies, a comprehensive sampling design is a key factor for not only improving inferential accuracy, but also for maximizing the explanatory power of the resulting phylogeny. Two standing problems in molecular phylogenies are the unstable placements of some deep and long branches, and the phylogenetic relationships shown by robust supported clades conflict with recognized knowledge. Empirical and theoretical studies suggest that increasing taxon sampling is expected to ameliorate, if not resolve, both problems; however, neither the current taxonomic system nor the established phylogeny provides sufficient information to guide additional sampling design. In the present study, we will select ingroup species for additional sampling based on epigynal morphology; reconstruct the molecular phylogeny of the micronetines-erigonines clade; interpret the phylogenetic relationships by tracing the evolution of epigynal characters; review the taxonomic system of micronetines based on the phylogenetic frame work. Our results provide a feasible criterion for high level phylogenetic inference with incomplete sampling to get a comprehensive representativeness; tracing the evolutionary changes of the epigynal characters with series and continuous transformations provide a efficient method to understand phylogenetic relationships with controversies.